The D780 features two autofocus systems depending on shooting mode. Through the optical viewfinder, it uses a 51-point phase-detection system. In live view mode, it switches to a hybrid on-sensor system combining phase and contrast detection with eye detection, face detection, and subject tracking capabilities. This dual approach gives you traditional DSLR speed when shooting through the viewfinder and modern tracking performance for video or tilted-screen work.

The D780 records 4K video at up to 30fps with 8-bit color depth. It offers both standard and flat color profiles for post-production flexibility. The tilting 3.2-inch touchscreen makes framing easier during video work, and dedicated headphone and microphone jacks (both 3.5mm) provide proper audio monitoring and input options. The hybrid autofocus system with eye and subject tracking works during video recording.

The D780 does not include in-body image stabilization. For stabilized shooting, you'll need to use Nikon lenses with built-in VR (Vibration Reduction) technology. This is common for DSLRs in this category, where lens-based stabilization has traditionally been the standard approach.

The D780 features dual SD card slots that accept SD, SDHC, and SDXC cards. This dual-slot design allows for backup recording, overflow storage, or separating RAW and JPEG files across cards for efficient workflow management.

The D780 features a weather-sealed magnesium alloy body designed to withstand dust and moisture. The weather sealing covers key seams and controls, making it suitable for outdoor shooting in challenging conditions when paired with weather-sealed lenses. The robust 840g magnesium construction balances durability with reasonable weight for all-day shooting.